The thresher shark is an all-encompassing term referring to three surviving species from the family Alopiidae: the pelagic thresher (Alopias pelagicus), the bigeye thresher (Alopias superciliosus), and the common thresher (Alopias vulpinus).
Thresher jaws are small. They have clearly not evolved to attack prey like tuna and seals; if anything, they look like they should be eating small crabs or scavenging chunks of floating debris that someone else took the time to kill. They would seemingly struggle to catch a live fish because the long snout overhangs their tiny mouth so much. That’s where the notorious tail comes in. The thresher tail is designed to be used like a whip to strike and immobilise prey , . Such force occurs with a single tail slap that gas is diffused out of the seawater and rises to the surface in bubbles . Typically, pelagic sharks hunt and capture one fish at a time; this strategy enables the shark to catch on average 3 fish, and sometimes more, in one go.
The easiest way to differentiate between the three species is the colouration. Common threshers are likely to be more grey and they lack any colouration above their pectoral fins; sometimes, they have white dots on the tips of their fins. Pelagic threshers have distinct colouration above their pectoral fins. The bigeye’s eye is visible from the top of the head and has characteristic groves above the eyes and gills.
Threshers are pelagic, meaning they live in the deep, open ocean. The only thresher known to inhabit New Zealand waters is the largest of the trio, the common thresher . The bigeye may also be found in NZ, but due to its habit of staying hundreds of metres below the ocean surface during the day, it is unlikely to be encountered if it is there at all .
The common and bigeye thresher are ‘Vulnerable’ to extinction under the IUCN Red List , , and the pelagic thresher is further threatened and classified as ‘Endangered’ . Estimated global population reductions are as follows:
Common thresher: 30–49% over 3 generations (76.5 years)
Bigeye thresher: 30–49% over 3 generations (55.5 years)
Pelagic thresher: 50–79% over 3 generations (55.5 years)
All three species are globally targeted for their meat, fins, skin, and liver oil , the latter commonly used in modern cosmetics and supplements. All species are usually retained if accidentally caught on commercial vessels –. In one study  conducted in the Indian Ocean, 13 out of 19 bigeye threshers caught on a commercial vessel’s longline were dead upon retrieval. In the same study, in the Atlantic Ocean, 412 out of 849 bigeye threshers caught on longlines were dead. On the same ships were deaths of stingrays and manta rays as well as blue, shortfin mako, silky, and smooth hammerhead sharks. One of the largest hubs for shark fin trading globally is Hong Kong , and threshers accounted for up to 3% of the total fin mass in the past .
Threshers are also popular among recreational, big game anglers. Although a tag and release method is more commonly practised nowadays, there is also a risk of post-release mortality. With threshers likely to be hooked from their tail due to their hunting style, a study of common threshers found 78% of tail-hooked sharks died after release .
The New Zealand commercial fishery is not exempt from thresher shark landings. A recent report by Fisheries New Zealand  saw that in 5 years from the 2014/15 to 2018/19 commercial fishing seasons, 149,916 tonnes of common thresher shark were caught by the core deep-water fleet. With the average 5-metre common thresher shark usually weighing in at 230 kg, we can infer that in the 5 years, the deep-water fleet landed approximately 651 individual thresher sharks.
Thresher vs swordfish
In April 2020, a dead, 4.5 metre, female bigeye thresher was found beached on a Libyan coast . A 30 cm swordfish (Xiphiasgladius) rostrum was found embedded in its head. It is understood that the rostrum severely injured some of the shark’s nerves, arteries, and gill arches. It was concluded that the impalement was what led to the ultimate death of the shark.
 S. P. Oliver, J. R. Turner, K. Gann, M. Silvosa, and T. D’Urban Jackson, “Thresher Sharks Use Tail-Slaps as a Hunting Strategy,” PLoS One, vol. 8, no. 7, 2013, doi: 10.1371/journal.pone.0067380.
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 C. L. Rigby et al., “Alopias superciliosus,” IUCN Red List Threat. Species 2019, vol. e.T161696A, 2019.
 R. W. Jabado et al., “The trade in sharks and their products in the United Arab Emirates,” Biol. Conserv., vol. 181, pp. 190–198, 2015, doi: 10.1016/j.biocon.2014.10.032.
 S. C. Clarke, J. E. Magnussen, D. L. Abercrombie, M. K. McAllister, and M. S. Shivji, “Identification of shark species composition and proportion in the Hong Kong shark fin market based on molecular genetics and trade records,” Conserv. Biol., vol. 20, no. 1, pp. 201–211, 2006, doi: 10.1111/j.1523-1739.2005.00247.x.
 A. T. Fields et al., “Species composition of the international shark fin trade assessed through a retail-market survey in Hong Kong,” Conserv. Biol., vol. 32, no. 2, pp. 376–389, 2018, doi: 10.1111/cobi.13043.
 S. C. Clarke et al., “Global estimates of shark catches using trade records from commercial markets,” Ecol. Lett., vol. 9, no. 10, pp. 1115–1126, 2006, doi: 10.1111/j.1461-0248.2006.00968.x.
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 Fisheries New Zealand, “Annual Review Report for Deepwater Fisheries for 2018/19,” 2020.
 P. L. Jambura, J. Türtscher, J. Kriwet, and S. A. A. Al Mabruk, “Deadly interaction between a swordfish Xiphias gladius and a bigeye thresher shark Alopias superciliosus,” Ichthyol. Res., vol. 68, no. 2, pp. 317–321, 2021, doi: 10.1007/s10228-020-00787-x.
 R. Coelho, J. Fernandez-Carvalho, P. G. Lino, and M. N. Santos, “An overview of the hooking mortality of elasmobranchs caught in a swordfish pelagic longline fishery in the Atlantic Ocean,” Aquat. Living Resour., vol. 25, no. 4, pp. 311–319, 2012, doi: 10.1051/alr/2012030.
The shortfin mako shark (Isurus oxyrinchus) is a pelagic, cartilaginous fish with a wide distribution range that covers most oceans and undergoes migrations that can be as large as 5,300km in just under 1.5 years (Barreto, de Farias, Andrade, Santana, & Lessa, 2016; Kohler, Turner, Hoey, Natanson, & Briggs, 2002). There is high demand for mako shark meat, and it is a prize game species in recreational fishing worldwide (Barreto et al., 2016).
As of 2019, I. oxyrinchus is classified as “Endangered” on the International Union for the Conservation of Nature’s (IUCN) Red List (Rigby et al., n.d.). Rigby et al. (n.d.) concluded that shortfin mako’s population trend is decreasing; there is an estimated decline everywhere except in the South Pacific and an overall estimated average reduction of 46.6% over 72-75 years. There is an estimated decline in biomass and abundance of 99.9% since the early 1800s, the main reason being overfishing (Ferretti et al., 2008).
Shortfin mako circulatory systems utilise a heat-exchanging technique that raises their internal temperature above that of the surrounding environment (Carey, Teal, & Kanwisher, 1981; Kohler et al., 2002). They have streamlined bodies, and aerobic muscles centred closer to their rear, which aids in thunniform swimming and increases power (Donley, Sepulveda, Konstantinidis, Gemballa, & Shadwick, 2004; Wegner, Sepulveda, Olson, Hyndman, & Graham, 2010). Emery and Szczepanski (1986) concluded that the gill area of I. oxyrinchus is 2-3 times larger than other pelagic shark species, which could aid in the mako’s speed, agility, and ability to swim long distances. Shortfin mako habitat extends globally in tropical and temperate oceans and they can be found inshore in coastal areas or at least 500m down in oceanic zones (Kohler et al., 2002).
Sexual dimorphism is prevalent in I. oxyrinchus,with females often occurring larger than males, with males reaching a maximum size of 2.6m and females reaching a maximum size of 3.4m (Barreto et al., 2016; Cema & Lincandeo, 2009; Chan, 2001; Doño, Montealegre-Quijano, Domingo, & Kinas, 2014; Hsu, 2001; Natanson et al., 2006; Semba, Nakana, & Aoki, 2009). Shortfin mako sharks are oophagous and ovoviviparous (Kohler et al., 2002), and a study by Mollet et al. (2000) predicts a gestation period of 15-18 months, although a study by Duffy and Francis (2001) puts makos in New Zealand waters at a 21-month gestation period. They have a 3-year reproductive cycle (Mollet & Cailliet, 2002). Bishop, Francis, Duffy, and Montgomery (2006) concluded that New Zealand shortfin mako births are concentrated in spring and gave a theoretical birth date of 1 October, with the average length of the shark at birth to be 61cm. Francis and Duffy’s (2005) study on sexual maturity of New Zealand shortfin makos concluded that maturity occurs between 7-9 years for males and 19-21 years for females (Bishop et al., 2006). Bishop et al. (2006) found evidence of New Zealand shortfin makos living to 29 years, although this number is probably higher because there is a lower chance of catching older sharks which make up a small percentage of the overall population. The same study found the sharks grow quickly within their first year after birth; this growth rate rapidly reduces in the subsequent years to steadier growth. Late maturity, moderately long longevity, the estimated low natural mortality rate, and low annual fecundity causes low productivity in the species (Bishop et al., 2006; Mollet et al., 2000).
Stillwell and Kohler (1982) analysed the stomach contents of shortfin mako sharks and found evidence of bony fish, swordfish, and cephalopods. As shark body length increased, so did the average volume of food, indicating that as makos grow larger, they may switch to larger prey items (Kohler et al., 2002).
Sharks have evolved for 400 million years (Donley et al., 2004), leading species such as I. oxyrinchus to be apex predators at the top of their food chain; they have no natural predators which results in low natural mortality. There is evidence of parasitic activity in shortfin makos, although it could not be determined if there was a negative effect on the shark (Borucinska & Hege, 1999). One example of anthropogenically influenced mortality is seen in 1966 when a longfin mako (Isurus paucus) died from a fishing hook retained in its flesh (Adams, Borucinska, Maillett, Whitburn, & Sander, 2015).
Sharks are keystone species and have a strong effect on multiple ecosystems due to their predatory role and wide dispersal range (Feretti, Worm, Britten, Heithaus, & Lotze, 2010). I. oxyrinchus is considered to be a large shark with “strong, top-down forces” (Feretti et al., 2010, p1055). Therefore, their removal from an ecosystem is highly likely to drastically alter communities, induce trophic cascades, release mesopredators such as smaller sharks and rays, and consequently cause a decline in commercial fish stocks (Fig. 1).
There is a high commercial demand for shortfin mako shark meat, and the species is exploited globally (Barreto et al., 2016). Peru is one of the biggest shark-fishing nations, and I. oxyrinchus is one of the top 2 most caught species with a rarely enforced catch size limit (Gonzalez-Pestana, Kouri, & Velez-Zuazo, 2016; Fischer, Erikstein, D’Offay, Guggisberg, & Barone, 2012). Brazilian fleets in the western and central South Atlantic exploit immature shortfin makos, specifically females (Barreto, 2016); it was rare for them to catch individuals greater than 2m. On a global scale, it is both a target and bycatch species in commercial and small-scale fisheries, including longline, gillnet, purse seine, trammel net, and trawls (Camhi, Pikitch, & Babcock, 2008; Rigby et al., 2019). There is likely underreporting of catch, and commercial post-release mortality from longlines alone is reported at 30-33% (Campana, Joyce, Fowler, & Showell, 2016; Rigby et al., 2019). Shortfin mako fins made up 1.2% of shark fin imported to Hong Kong in 2014 (Fields et al., 2017), and their skin, jaws, and liver oil are also used (Compagno, 2001). In New Zealand’s EEZ, I. oxyrinchus is a common bycatch species on tuna longlines and less commonly on pelagic longlines, trawls, and set nets (Bagley et al., 2000; Francis, 1998; Francis, Griggs, Baird, Murray, & Dean, 2000; Martinsohn & Muller, 1992). Since 1993, commercial catch averaged 60 tonnes annually, but observer reporting estimated 100-200 tonnes per year from the longline tuna fishery alone (Francis, 1998; Francis et al. 2000); the discrepancy between these numbers could be related to lack of accurate recording. From 1988-2015, the New Zealand tuna longline fishery total catch was comprised of 11.1% target species (southern bluefin tuna, bigeye tuna, and swordfish) and 88.9% bycatch (including albacore tuna, lancetfish, porbeagle shark, deepwater dogfish, dealfish, mako shark, moonfish, escolar, sunfish, and butterfly tuna) according to a report by Fisheries New Zealand [FNZ] (2018). Since October 2014, shark finning is illegal in New Zealand, yet over half of all makos caught by charter vessels in the 2014-15 tuna longline fishery year were kept for their flesh (FNZ, 2018). The low productivity of mako sharks causes them to be vulnerable to overfishing and makes it hard for the species to replace lost individuals. CITES (2019) reported that shortfin mako is in danger of population collapse due to the overfishing of most of the juveniles since the 1980s; this means that sharks dying of old age now will not be replaced with mature individuals leaving a 10-20-year gap.
The mining of precious metals increases the presence of toxic contaminants in the environment, especially in coastal environments, specifically mercury (Maz-Courrau et al., 2011). Sharks bioaccumulate mercury through their tissues and organs from the environment and through the food they eat; mercury travels up the trophic web and magnifies at each level, eventually accumulating in shark muscular tissue (Maz-Courrau et al., 2011). Maz-Courrau et al. (2011) found that 33% of shortfin mako sharks they studied off the Baja California coast had levels of mercury higher than was estimated to be fit for human consumption by Mexican Law and Watling et al. (1981) found mercury concentrations in mako sharks off the coast of Australia to be almost twice that. As Maz-Corrau et al. (2011) only sampled juveniles, it is likely that as sharks grow and eat more, more mercury is accumulated into their tissue over time. When people eat this contaminated tissue, it bioaccumulates up the food chain again and causes adverse health effects in humans.
Current/Proposed Management Actions
As of 2019, I. oxyrinchus officially met criteria to be listed on the Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES) Appendix II, which “includes species not necessarily threatened with extinction, but in which trade must be controlled in order to avoid utilisation incompatible with their survival” (CITES, 2019, p. 2).
Pressure from overfishing appears to be the most significant contributor to shortfin mako shark decline globally. Due to the widespread distribution of I. oxyrinchus and migratory nature, conservation effort and management is needed at both a local and international scale (Adams, Flores, Flores, Aarestrup, & Svendsen, 2016; Corrigan et al., 2018). Because of their “Endangered” status on the with a decreasing population trend, it is essential to increase their population trend and bring them to the next highest, more stable level on the Red List, “Vulnerable”, and take measures to ensure they do not go the other direction and become classified as “Critically Endangered”. The most effective way this can be achieved is through strict management of fisheries and the implementation of Marine Protected Areas (MPA) in habitats that are suitable for shortfin makos by means of international treaties (Birkmanis, Partridge, Simmons, Heupel, & Sequeira, 2020; Dulvy et al., 2008; Rigby et al., 2019).
Community Education & Awareness
Films, documentaries, and TV series are a great way of moving an audience and opening their eyes to what is going on in the world around them in an entertaining way. Saving Jaws by Ocean Ramsey, Shark Week by The Discovery Channel, and BBC’s Blue Planet, narrated by David Attenborough, are a few.
For the booklovers, don’t worry, we got you. Ocean Ramsey’s What You Should Know About Sharks: Shark Language, social behavior, human inter-actions, and life saving information educates readers on shark behaviours, how to swim with them safely, and debunks common shark myths. Sylvia A. Earle’s book The World is Blue: How Our Fate and the Ocean’s are One shows readers how every one of us literally breathes the ocean, and her other book, Sea Change: A Message of the Oceans, has been compared to Rachel Carson’s Silent Spring. For those who do not enjoy reading, Christian Vizl’s Silent Kingdom is a photography-based book that shows the raw beauty, power, and elegance of marine creatures, such as sharks, in stunning black and white photos.
NGO’s also play an essential role in community education and awareness. In New Zealand, groups such as Auckland Whale & Dolphin Safari and Whale Watch Kaikōura take their customers to visit sea life up close. SEA LIFE Kelly Tarlton’s Aquarium educates both adults and children alike and cares for sick and dying turtles.
Monitoring & Research
There is enough research on I. oxyrinchus populations to know something has to change, but what lacks is monitoring the factors contributing to their dismal fate. As of 2019, CITES began to monitor the commercial catch and trade of shortfin mako but agreed that it might not be enough (CITES, 2019). In 2013, MPI released a National Plan of Action for the Conservation and Management of Sharks (NPOA). In the plan, they stated one of their objectives was to “systematically review management categories and protection status to ensure they are appropriate to the status of individual shark species” (MPI, 2013, p. 19) among others, yet there is a lack of reporting available for public access to back this up. They also had intentions of issuing a revised NPOA in 2018, which seems not to exist altogether.
Bycatch records need to be scrutinised and fisheries must be forced to show accurate record-keeping. Data collected from accurate bycatch records of I. oxyrinchus can then be accumulated around the globe to highlight the fisheries that are exploiting shortfin makos.
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